This figure shows the tail shape developed by three tadpole genotypes (M, N, and O) when raised in an environment with predators and an environment without predators. In the presence of predators, tadpoles with large tails have high fitness and tadpoles with small tails have low fitness. When predators are not present, tadpoles with large tails have low fitness and tadpoles with small tails have high fitness. 
Which genotype(s) would have the highest average fitness across both environments?
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