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Females Heterozygous for the Recessive Second Chromosome Mutations Px, Sp

question 68

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Females heterozygous for the recessive second chromosome mutations px, sp, and cn are mated to a male homozygous for all three mutations.The offspring are as follows:
\begin{array}{|l|r|}\hline p x  s p  c n & 1461 \\\hline p x  s p  c n^{+} & 3497 \\\hline p x  s p{+ c n} & 1\\\hline p x  s p^{+}  c n+ & 11 \\\hline p x^{+}  s p  c n & 9 \\\hline p x^{+}  s p  c n+ & 0 \\\hline p x^{+}  s p+ c n & 3483 \\\hline p x^{+}  s p+ c n+ & 1539 \\\hline & 10,000 \\\hline\end{array}
-Which of the three genes is in the middle?


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