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Healthy individuals of Paramecium bursaria contain photosynthetic algal endosymbionts of the genus Chlorella. When within their hosts, the algae are referred to as zoochlorellae. In aquaria with light coming from only one side, P. bursaria gather at the well-lit side, whereas other species of Paramecium gather at the opposite side. The zoochlorellae provide their hosts with glucose and oxygen, and P. bursaria provides its zoochlorellae with protection and motility. P. bursaria can lose its zoochlorellae in two ways:
(1) if kept in darkness, the algae will die; and
(2) if prey items (mostly bacteria) are absent from its habitat, P. bursaria will digest its zoochlorellae.
Which term best describes the symbiotic relationship of well-fed P. bursaria to their zoochlorellae?


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