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Dragonfly species A (with a carrying capacity of 1,000) and species B (with a carrying capacity of 400) are competing. The competition coefficient of species A on species B is 0.25, and the competition coefficient of species B on species A is 0.5. Dragonfly species C (with a carrying capacity of 5,000) and species D (with a carrying capacity of 10,000) are also competing. The competition coefficient of species C on species D is 0.1, and the competition coefficient of species D on species C is 0.2. Based on the Lotka‒Volterra equation, which of these pairs of species can coexist?
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