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The Substitution Rate Is Expressed in Terms of Substitutions Per

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The substitution rate is expressed in terms of substitutions per generation; however,molecular clock estimates show that substitutions accumulate annually regardless of the generation time.What explanation for this discrepancy is illustrated in the figure below?
The substitution rate is expressed in terms of substitutions per generation; however,molecular clock estimates show that substitutions accumulate annually regardless of the generation time.What explanation for this discrepancy is illustrated in the figure below?


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