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Performing a Complementation Test Through Cotransduction Using Mutant Phage Genomes

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Essay

Performing a complementation test through cotransduction using mutant phage genomes created by X-ray mutagenesis (mutants a-e),as well as ones created by chemical mutagenesis using EMS (mutants 1-5),you get the following results:
12345a++b+c+++d+++e++\begin{array} { l l l l l l } & 1 & 2 & 3 & 4 & 5 \\a & - & + & + & - & - \\b & - & + & - & - & - \\c & + & + & - & - & + \\d & + & - & + & + & - \\e & - & - & + & + & -\end{array}
Explain this data.

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