What is the nucleophilic species involved in the initial step of the reaction below? 
Definitions:
Linnaean Categories
Hierarchical levels of classification in biology developed by Carl Linnaeus, including kingdom, phylum, class, order, family, genus, and species.
Hierarchical
Describing a system or organization in which people or groups are ranked one above the other according to status or authority.
Domain
(1) A structural and functional region of a protein. (2) The broadest taxonomic category; each domain includes one or more kingdoms.
Paraphyletic Group
A group of organisms that includes an ancestral species and some, but not all, of its descendants.
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